Fallen

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The Fallen, also known as "spider pirates," are one of the enemy factions in the game.

The Fallen have only been seen present in Old Russia on Earth during the E3 Gameplay Reveal, but a quote on the Bungie site states that they have "...claimed every planet and moon Humans ever touched.", which could indicate they will be fairly pervasive throughout the game. The Fallen appear to be organized largely into groups called "Houses", seeming similar to clans or other family groups, which are not always friendly towards each other, and hunt in packs led by a "Fallen Captain".

In a video you can also see the Fallen teaming up with the Vex to attack the Guardians who just happened to cross the path of them both at the same time. We have also seen flying robots attack the players with the fallen in Old Russia. These may be of Fallen make or another species but seeing as only the fallen were present it is safe to assume they are of Fallen origin

Fallen Ranks:

Dreg:

"These relentless thieves seem driven by desperation and rage. They are crafty, but they usually give up their cover with their clattering and hissing. Most seem outfitted with some kind of shock dagger and pistol, and as far as we can gather, they seem to only have two arms, while other Fallen have four." -- Bungie Site (Our Enemies)

Vandal:

"The truths about the Fallen chain-of-command are still making themselves known, but Vandals appear to hold a higher ranking than the Dregs, and deadlier weapons, too. They often have long range capability, but, lucky for us, their weapons take time to charge." -- Bungie Site (Our Enemies)

Captain:

A confirmed Captain rank Fallen is Rixis.

"Fallen run in packs, led by an individual Captain. There is no denying a Captain's might; they are the fiercest of the Fallen. There is this sense of tattered nobility to them. You can see it in the way they carry themselves. They think they're better than us. They believe that Earth belongs to them." -- Bungie Site (Our Enemies)

Speculation:

"...there’s just something wonderful about the idea of a nomadic race of once noble houses now scattered to the solar winds." -- Urk (March of the Mail Sack, Destiny News)

Many developers have alluded to the Fallen Houses having, well, fallen, however, not much beyond teasers is known on this point.